Elpošanas ķēdes loma piruvātu producējošos Zymomonas mobilis celmos

6533b852fe1ef96bd12aa711

RESEARCH PRODUCT

Elpošanas ķēdes loma piruvātu producējošos Zymomonas mobilis celmos

subject

elpošanas ķēdes mutants Zymomonas mobilis elektronu-transporta ķēde Bioloģija piruvātdekarboksilāze laktātdehidrogenāze

description

Lai novirzītu Zymomonas mobilis katabolismu no etanola producēšanas uz piruvāta producēšanu, jāatrod veids, kā oksidēt to NADH, kas parasti tiek izmantots etanola sintēzei, bet nav nepieciešams piruvāta producēšanai. Elpošana ir viens veidiem, kā oksidēt NADH. Lai gūtu priekšstatu par elpošanas ķēdes ietekmi uz piruvāta akumulēšanos Z. mobilis, tika salīdzināti vairāki elpošanas ķēdes mutantie celmi. Uz daļēji inaktivētas piruvātdekarboksilāzes (PDC) fona darbā izdevās iegūt celmu ar pārekspresētu membrānsaistīto D-laktātdehidrogenāzi (LDH), bet neizdevās iegūt celmu ar pārekspresētu NADH dehidrogenāzi. Novērojām, ka elpošanas ķēdes NADH dehidrogenāzes un LDH aktivitātes aerobi augošās Z.mobilis šūnās nesummējas - celmā ar pārekspresētu LDH aktivitāti, bet aktīvu NADH dehidrogenāzi, skābekļa patēriņš notiek ar līdzīgu vai pat mazāku ātrumu nekā celmā Zm6, kurā LDH nav pārekspresēta. Noskaidrojām, ka celmos ar daļēji inaktivētu piruvātdekarboksilāzi elpošana nav limitējošais faktors piruvāta akumulācijā, jo līdzīga piruvāta akumulācija notiek pie dažādiem elpošanas ātrumiem. Lai gan celmiem ar daļēju PDC inaktivāciju ir novērojams 50% kritums tās aktivitātē, lielākā daļa glikozes oglekļa joprojām nonāk etanolā un CO2.

year	journal	country	edition	language
2018-01-01

https://dspace.lu.lv/dspace/handle/7/38634

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