Palaeoenvironmental andstratigraphic aspects of animal traces and plant remains in Spanish Permian red beds (Peña Sagra, Cantabrian Mountains, Spain)

Abstract Various vertebrate and invertebrate traces and plants remains have been discovered in the Permian of Pico Paraes (Cantabrian Mountains). They occur in the Sagra Formation, a 400 m thick unit consisting of a vertical alternation of channels with conglomeratic sandstones and fossiliferous red siltstones with a caliche horizon at the top. The most common trace is Hyloidichnus major which is attributed to approximately 50 cm long Captorhinomorphs. Other tracks provisionally assigned to Limnopus could also indicate the presence of Temnospondyls. The bedding planes with traces also show some tracks of Isopodichnus cf. minutus which is ascribed to small triopsid crustaceans. Like the foot…