0000000000985296

AUTHOR

Tadeusz Zatwarnicki

<p><strong>A revision of the Old World species of <em>Leptopsilopa</em> Cresson (Insecta: Diptera: Ephydridae)</strong></p>

Diagnoses and illustrations of three Leptopsilopa species known in the Old World: L. africana Cresson, L. mutabilis Cresson and L. pollinosa (Kertesz) are provided. Three new synonyms are proposed: L. aurata (Canzoneri and Meneghini) and L. tibialis (Canzoneri and Meneghini) = L. africana; Psilopa confeteae Canzoneri and Munari = L. pollinosa . Three other species are removed from Leptopsilopa and combined with Athyroglossa , as A. leonensis (Canzoneri and Rampini), new combination, A. mianii (Canzoneri) new combination and A. rossii (Canzoneri and Raffone), new combination.

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Notes on Shore Flies (Diptera: Ephydridae) from Finland and north-western Russia

The recent checklist of the Ephydridae of Finland by Zatwarnicki and Kahanpää (2014) mentioned 13 ephydrid species as new for Finland without further details. This paper presents detailed records for those species and a few other species of interest. Four species are recorded for the first time from Russia. Trimerina indistincta Krivosheina, 2004 is herein considered as a new junior synonym of Trimerina microchaeta Hendel, 1932, syn. nov.

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A Revised Phylogeny of Discocerinini (Diptera: Ephydridae) with an Emphasis on Structures of the Proboscis

Supplementary diagnostic characters of genera in the tribe Discocerinini are suggested. A new genus Aquachasma Zatwarnicki, type species: Discocerina leucoprocta Loew, 1861 is described with 24 new combinations of all included species. Genus Polytrichophora is divided into the nominate subgenus and a new subgenus Sklodowskopa Zatwarnicki, type species: Clasiopa duplosetosa Becker, 1896. Genus Orasiopa is divided into the nominate subgenus and a new subgenus Reymontopa Zatwarnicki, type species: Discocerina mera Cresson, 1939. The shape of the proboscis of all genera of Discocerinini is provided, including variability of cibarium and pseudotracheae. The ventral receptacles of 6 genera (Aquac…

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A revision of the new world species of Polytrichophora Cresson and Facitrichophora, new genus (Diptera, Ephydridae)

The New World species of Polytrichophora Cresson and Fascitrichophora new genus, are revised. Fifteen new species are described (type locality in parenthesis): Fascitrishophora atrella sp. n. (Costa Rica. Guanacaste: Murciélago [10°56.9’N, 85°42.5’W; sandy mud flats around mangrove inlet]), F. carvalhorum sp.n. (Brazil. São Paulo: Praia Puruba [23°21’S, 44°55.6’W; beach]), F. manza sp. n. (Trinidad and Tobago. Trinidad. St. Andrew: Lower Manzanilla (12 km S; 10°24.5’N, 61°01.5’W), bridge over Nariva River), F. panama sp. n. (Panama. Darien: Garachine [8°04’N, 78°22’W]), Polytrichophora adarca sp. n…

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Catalogue of the Egyptian Ephydroidea (Diptera: Schizophora: Acalyptratae)

We present a catalogue of all known taxa of the superfamily Ephydroidea in Egypt. Old World synonymies, type localities, type depositories, world distributions by biogeographic realm(s) and country, Egyptian localities and dates of collection are provided. A total number of 117 species belonging to 45 genera, 18 tribes, 7 subfamilies, and representing 3 families has been catalogued. The treated families are: Ephydridae (shore flies or brine flies), Drosophilidae (vinegar flies or fruit flies) and Braulidae (bee lice). One shore fly species, Psilopa clara (Wollaston, 1858), is recorded for the first time from Egypt. Two new synonyms of shore flies are proposed, namely: Philotelma ulianai Raf…

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A Revision of the Nearctic Species of Hecamedoides Hendel (Diptera: Ephydridae)

Abstract Species of the shore-fly genus Hecamedoides Hendel from the Nearctic Region are revised, including description of H. lattini, new species (Virginia. Stafford: Falmouth (38°19.2′N, 77°28.1′W; Rappahannock River; 9 m). To provide context and also to facilitate identification, diagnoses are provided for the tribe Discocerinini and genus in addition to a key to the genera and species, H. lattini and H. unispinosus, occurring in the New World. Diagnostic characters, especially of the male terminalia, are illustrated, and distribution maps are also provided.

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Fauna Europaea: Diptera – Brachycera

Fauna Europaea provides a public web-service with an index of scientific names (including important synonyms) of all extant multicellular European terrestrial and freshwater animals and their geographical distribution at the level of countries and major islands (east of the Urals and excluding the Caucasus region). The Fauna Europaea project comprises about 230,000 taxonomic names, including 130,000 accepted species and 14,000 accepted subspecies, which is much more than the originally projected number of 100,000 species. Fauna Europaea represents a huge effort by more than 400 contributing taxonomic specialists throughout Europe and is a unique (standard) reference suitable for many user c…

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Heterogeneity in shore flies – the case ofGlenantheHaliday (Diptera: Ephydridae) in the Old World

Abstract Old World species of the genus Glenanthe are treated comprehensively with an emphasis on the unusually diverse morphological heterogeneity discovered in structures of the male terminalia. As perspective for this treatment, the tribe Lipochaetini, in which Glenanthe is placed, is characterized and discussed, and an annotated key to the four included genera is provided to facilitate their identification. Glenanthe is demonstrated to be a monophyletic lineage within Lipochaetini. Seven Old World species of Glenanthe are now known, including two new Afrotropical species as follows (type locality in parenthesis): G. namibia n. sp. (Namibia. Mukwe: Divuju: Okavango River [18°04’04”S, 21°…

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New taxonomic twists in shore flies as revealed in discovery of Neozealides, a new genus from New Zealand (Diptera: Ephydridae)

We describe Neozealides to accommodate two endemic species from New Zealand that were previously placed in the shore-fly genus Hyadina Haliday. Although similar to Hyadina, both species are more closely related to the Hyadina and Philygria groups of genera (Zatwarnicki & Ryczko 2014) than they are to species within Hyadina. To document these discoveries, we revise both species, placing them in Neozealides. Emphasis is given to structures of the male terminalia, which are fully illustrated. Detailed locality data and distribution maps for both species are provided. For perspective and to facilitate recognition of genera, the tribe Hyadinini is diagnosed and a key to genera from New Zeala…

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Systematic Position of Gastropini (Diptera: Ephydridae), and a Review of Two Included Genera:GastropsWilliston andNotacanthinaMacquart

The placement of the tribe Gastropini within the subfamily Gymnomyzinae, with the tribe Gymnomyzini as a sister-group, is supported. Validity for two genera Gastrops Williston and Notacanthina Macquart (senior synonym of Beckeriella Williston) is maintained. Particular characters of their proboscis, ventral receptacle and the male terminalia are presented. A key to species and habitus photographs of selected species of Gastrops and Notacanthina are provided. The following new combinations are proposed: Notacanthina clypeata (Lizarralde de Grosso, 1986), comb. nov., N. fasciata (Mathis & Grimaldi, 2000), comb. nov., N. filipina (Lizarralde de Grosso, 1994), comb. nov., N. gigas (Lizarralde d…

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Notes on Selected Genera of Shore Flies (Diptera: Ephydridae), with Removals to Drosophilidae, Heleomyzidae and Milichiidae

The following new taxonomic changes in nine tribes of Ephydridae are proposed: status at species level: Ditrichophora amoena (Duda, 1942); 2 resurrections from synonymy: Ptilomyia madeirensis Stuke, 2012, bona species, and P. orsovana (Enderlein, 1922), bona species; 10 new combinations: Allotrichoma communis (Malloch, 1934), comb. nov., Athyroglossa bettelai (Canzoneri, 1995), comb. nov., Ceropsilopa africana (Canzoneri & Meneghini, 1969), comb. nov., Ceropsilopa flavescens (Frey, 1958), comb. nov., Diclasiopa affinis (Tonnoir & Malloch, 1926), comb. nov., Diclasiopa flavitarsis (Tonnoir & Malloch, 1926), comb. nov., Diclasiopa senegalensis (Canzoneri, 1981), comb. nov., Ditrichophora amoe…

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Review of the shore-fly genus Oedenopiforma (Diptera: Ephydridae)

The shore-fly genus Oedenopiforma Cogan is reviewed, including new information on Oedenopiforma argentea (distribution), Oedenopiforma javana (comb. nov. and O. orientalis syn. nov.), and Oedenopiforma vockerothi (sp. nov. from United Arab Emirates). Structures of the male terminalia of these three species, a diagnosis of the genus, and a key to the included genera of Dryxini and species of Oedenopiforma are presented. Resume´—Nous revisons le genre Oedenopiforma Cogan chez les ephydrides et apportons de nouvelles informations sur Oedenopiforma argentea (repartition), Oedenopiforma javana (nouvelle combinaison et O. orientalis nouvelle synonymie )e tOedenopiforma vockerothi (nouvelle espece…

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p class="ZootaxaTitle"First description of immature stages of Gastropini, Gastrops willistoni Cresson (Diptera: Ephydridae) preying on eggs in a foam nest of Leptodactylus knudseni Heyer (Anura: Leptodactylidae) in Central Amazon.

The third instar larva and puparium of Gastrops willistoni Cresson (Diptera: Ephydridae), preying on eggs in a foam nest of Leptodactylus knudseni Heyer (Anura: Leptodactylidae), in a central Amazonian dryland forest (Brazil) are described for the first time. The fusiform larva has a short breathing tube, its anterior spiracles are fan-shaped and posterior spiracles with three spircular openings. Pupa is ovoid with a broad anterior margin and bifurcate posterior tip. The taxonomic relationship of the frog egg predators, Gastrops Williston and Typopsilopa Cresson, are discussed. 

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A Revision of theniveaGroup of the Shore-Fly GenusDitrichophoraCresson (Diptera: Ephydridae)

ABSTRACT Species of the nivea group (genus Ditrichophora Cresson) are revised. The group now includes four species of which one, D. gulkana (Alaska and Mongolia), is newly described. The group is now known to have a Holarctic distribution with two species, D. gulkana and D. nivea, being the first reported from the Nearctic Region. Another Asian species, D. brunnicosa, previously placed in Diclasiopa, is transferred to this group (genus Ditrichophora) and is also reported from Mongolia. To facilitate identification of these rare species, we have included diagnoses of the tribe, genus, and species group and have also provided an annotated key to Holarctic genera of the tribe Discocerinini and…

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Bibliografia Dipterologiczna Polski (1597-2020). Polish Dipterological Bibliography (1597-2020)

An outline of the history of Diptera research and the achievements of dipterology in Poland is described. A list of 4470 publications on Polish Diptera and papers, book chapters, and books of Polish dipterists that have appeared from 1597 is provided. The list was closed on 31. Dec. 2020

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A revision of the new world species of Gymnoclasiopa Hendel (Diptera, Ephydridae)

Species of the shore-fly genus Gymnoclasiopa Hendel from the New World are revised, including G. grecorum, sp. n. (Alaska. Juneau: Gastineau Channel, Thane Road (S Juneau; 58°16.9’N, 134°22.4’W)) and G. matanuska, sp. n. (Alaska. Matanuska-Susitna: Palmer (Matanuska River; 61°36.5’N, 149°04.1’W)). We also clarify the status of previously described species, including those now discovered to have Holarctic distributions and/or for which sexual dimorphism was not appreciated and the species was described twice, including G. montana (Cresson) as a syn. n. of G. bohemanni (Becker). Two species, G. bella (Mathis), comb. n., and G. chiapas (Mathis), comb. n., are transferred from Ditrichophora to …

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A revision of the shore-fly genus Hydrochasma Hendel (Diptera, Ephydridae)

A revision of the shore-fly genus Hydrochasma Hendel. The species of the genus Hydrochasma Hendel are revised, including 27 new species (type locality in parenthesis): H. andeum (Ecuador. Guayas: Boliche (02°07.7'S, 79°35.5'W)), H. annae (United States. Utah. Grand: Swasey Beach (15.3 km N Green River; 39°07'N, 110°06.6'W; Green River; 1255 m)), H. capsum (Ecuador. Orellana: RíoTiputini (0°38.2'S, 76°8.9'W)), H. castilloi (Ecuador. Loja: Catamayo (03°59'S, 79°21'W)), H. crenulum (Peru. Cuzco: Paucartambo, Atalaya (Río Alto Madre de Dios; 12°53.3'S, 71°21.6'W; 600 m)), H. denticum (Ecuador…

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Checklist of the family Ephydridae of Finland (Insecta, Diptera)

A checklist of 112 species of shore flies (Ephydridae, Diptera) recorded from Finland is presented. Comparing this to the list of Hackman (1980), 52 changes are made: 25 species are added (all but one recorded after 1980), 18 misidentifications are deleted, 5 junior synonyms are replaced and 5 updated generic combinations are given. Peer reviewed

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Taxonomic notes on the genera Scatella and Scatophila (Diptera: Ephydridae) with a remark on Trixoscelis chilensis (Trixoscelididae).

Two new combinations are proposed in Scatella for Ephydra australis Walker, 1853 and Opomyza guttata Macquart, 1843. Two species are resurrected from synonymy: Scatella lacustris (Meigen, 1830), revised status and Trixoscelis chilensis (Schiner, 1868), revised status (Trixoscelididae). Nine new synonyms are suggested: Scatella insularis Mathis & Wirth, 1981 = S. australis (Walker, 1853); S. lutosa nigripes Oldenberg, 1923 = S. obsoleta Loew, 1861; S. septempunctata Malloch, 1933 = S. gratiellae Canzoneri & Raffone, 1987 = S. septemfenestrata Lamb, 1912; S. tenuicosta Collin, 1930 = S. lacustris (Meigen, 1830); S. vulgata Cresson, 1931 = S. guttata (Macquart, 1843); Scatophila zlobin…

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Revision of the Shore-fly Genera Parahyadina Tonnoir and Malloch and New Zealand Hyadina Haliday (Diptera: Ephydridae).

Two genera of the shore-fly tribe Hyadinini, Hyadina Haliday and Parahyadina Tonnoir & Malloch, are revised, and all species treated, save P. lacustris Tonnoir & Malloch, are described for the first time. The phylogenetic position of both genera within Hyadinini is presented along with confirming, morphological evidence. The New Zealand fauna of Hyadina is now limited to the single new species H. breva, and Parahyadina is expanded from being a monotypic genus based on P. lacustris to also include the following 9 endemic new species: P. angusta, P. atra, P. bifurcata, P. bulla, P. debilis, P. edmistoni, P. hennigi, P. irwini, and P. latistylis. For both genera, emphasis is given to s…

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Solving the Puzzle of Taxonomic Position of the Petroleum Fly by Resurrection of Diasemocera Bezzi from Psilopa Fallén (Diptera: Ephydridae) with Proposed Specific and Generic Synonymies

Diasemocera Bezzi, 1895 (Type species: Psilopa nigrotaeniata Bezzi, 1895 [= Psilopa roederi Girschner, 1889]) is resurrected from synonymy with Psilopa and the following four new synonyms of Diasemocera are proposed: Domina Hutton (Type species: Domina metallica Hutton), Discocerinella Mercier (Type species: Discocerinella omonvillea Mercier, 1927 [= Notiphila pulicaria Haliday, 1839]), Trimerinoides Cresson (Type species: Trimerina adfinis Cresson) and Helaeomyia Cresson, 1941 (Type species: Psilopa petrolei Coquillett). Diasemocera is placed close to the genus Trimerina and it differs from the highly specialized Psilopa and related genera by the number of pseudotracheae (6–7), vestiture o…

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40 lat minęło. Rubinowy jubileusz Sekcji Dipterologicznej Polskiego Towarzystwa Entomologicznego

40 years have passed. Ruby jubilee of the Dipterological Section of the Polish Entomological Society. The illustrated notes on the Dipterological Section of the Polish Entomological Society are presented. The idea of establishing the Section emerged in 1980 in Cracow, formally implemented a year later. Each year members meet at a scientific symposium in various places of Poland. The group releases international journal Dipteron, and in 2013 proclaimed World Dipteran Day.

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New discoveries in New World Atissini as revealed in revision of Pelignellus Sturtevant & Wheeler (Diptera: Ephydridae)

Pelignellus is resurrected from synonymy with Atissa and is revised. The genus and both included species—P. subnudus Sturtevant & Wheeler from the USA (California) and P. freidbergi n. sp. from Central America (Costa Rica, El Salvador, Panama)—are described with an emphasis on structures of the male terminalia, which are fully illustrated. The species are keyed and their distribution data are provided. For perspective and to facilitate recognition of genera and species, the tribe Atissini is diagnosed and a key to genera is provided.

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Revision of New World Species of the Shore-fly Subgenus Allotrichoma Becker of the Genus Allotrichoma with Description of the Subgenus Neotrichoma (Diptera, Ephydridae, Hecamedini)

The New World species of the subgenera Allotrichoma Becker and Neotrichoma (new subgenus) are revised, including a phylogenetic analysis of the species groups and subgenera within the genus Allotrichoma. For phylogenetic perspective and to document the monophyly of the genus Allotrichoma and its included subgenera and species groups, we also provide a cladistic analysis of genera within the tribe Hecamedini. The ingroup included seven exemplar congeners from within Allotrichoma. Outgroup sampling included exemplars of other genera within Hecamedini and from the putative sister group, Lipochaetini, and to root the analysis, we used an exemplar of the tribe Discocerinini. Analyses with succes…

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An overview of the Ephydridae (Diptera) of Saudi Arabia

Despite the species richness of Ephydridae world-wide (2000 species) and its prominent environmental roles as a minor pest and as a food for wildlife, only 13 species have been recorded from Saudi Arabia. Between 2012 and 2016, a biodiversity study of Diptera was conducted at Jazan, Asir, and Najran in south-western Saudi Arabia, at 22 sites, was performed mainly using Malaise traps and sweep nets. In this study, 43 known species of Ephydridae were identified, 37 of them for the first time from southwestern Saudi Arabia and 16 from Arabian Peninsula. This brings the total number of Ephydridae species in Saudi Arabia to 49 (including previous records). There were a further four species, whic…

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First Nearctic Record of Hydrellia albilabris (Meigen) (Diptera: Ephydridae), a Leafminer of Duckweed (Araceae: Lemnoideae), with Comments on Related Species

Hydrellia albilabris (Meigen) (Diptera: Ephydridae) is recorded in the Nearctic Region for the first time. Its biology and immature stages are documented, and other miners in duckweeds (Araceae: Lemnoideae) are discussed. One new combination is proposed: Hydrellia jinpingensis (Zhang, Yang, and Hayashi) is transferred from Cavatorella.

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Phylogeny and Taxonomy of the Risa Genus-group (Diptera: Ephydridae), with Description of a New Genus from Australia

The systematic and somewhat controversial history of Risa Becker is presented, and its relationship with Diasemocera Bezzi (tribe Psilopini, Ephydridae) is documented by morphological evidence and an association with host plants in the family Amaranthaceae. The tribe Risini Papp (as Risidae) is synonymized with Psilopini Cresson. Notorisa gen. nov., from Australia, is described (type species: Notorisa mcalpinei sp. nov.; Australia. Victoria: Big Desert National Park, near Lake Hindmarsh; 36°03.7'S 141°54.8'E). Achaetorisa Papp is retained as a subgenus within Risa and includes five species, including two new combinations: Risa brevicornis (Papp) comb. nov., Risa salsolae (Mathis & Zatwa…

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Phylogeny of Hyadinini (Diptera: Ephydridae) with an Emphasis on Structures of the Proboscis

Abstract. The shape of adult mouthparts (proboscis) of all genera of Hyadinini (Diptera: Ephydridae) is provided, including variability of cibarium, lacinia and the number of pseudotracheae. Its usage in phylogenetic construction is documented. All ten genera are diagnosed, including the genus Lytogaster, which is formally restored from synonymy with Hyadina. The ventral receptacle of four genera (Garifuna, Parahyadina, Parydroptera and Pelinoides) and the male terminalia of Parahyadina are presented for the first time. Monophyly of the tribe is discussed and the relationships among genera of Hyadinini are proposed. Ten Hyadinini genera are grouped into four lineages 1) Pelina group with Pe…

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Biogeographic Anomalies in Shore Flies as Revealed in Revision of the Shore-Fly Genus Subpelignus Papp (Diptera: Ephydridae)

Subpelignus Papp is revised, including description of (1) a new species, S. gagnei Zatwarnicki and Mathis (New Zealand. North Island. ND: Whananaki South (35°31.1'S, 174°27.2'E); (2) two new combinations, Subpelignus antennalis (Aldrich, 1931) and Subpelignus limosinus (Becker, 1896); (3) a new synonym, S. hortobagensis Papp, 1983 = S. limosinus Becker; and (4) description of a new subgenus, Niratissa Zatwarnicki and Mathis, for S. maculipennis Mathis and Zatwarnicki. Descriptions of revised and the newly described species include some of the first illustrations of structures of the male terminalia. Detailed locality data for all species examined are provided. For perspective and to facilit…

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<p class="HeadingRunIn"><strong>A review of the genus <em>Discomyza </em>(Diptera: Ephydridae) from Afrotropical, Australasian/Oceanian and Oriental Regions</strong></p>

Tropical Old World species of the genus Discomyza are redescribed, including male terminalia, photographs of heads and wings. One new species Discomyza fagomoga sp. nov. is described from D. R. Congo, Ethiopia, Kenya and Uganda. Two new synonyms are proposed: Discomyza obscuricornis Canzoneri and Rampini, 1996 = D. dolichocerus Cresson, 1944 and D. intermedia Canzoneri and Meneghini, 1969 = D. eritrea Cresson, 1939. A key to all species is provided. Relationships among species of Discomyza are suggested.

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Catalogue of the Diptera (Insecta) of Morocco— an annotated checklist, with distributions and a bibliography

The faunistic knowledge of the Diptera of Morocco recorded from 1787 to 2021 is summarized and updated in this first catalogue of Moroccan Diptera species. A total of 3057 species, classified into 948 genera and 93 families (21 Nematocera and 72 Brachycera), are listed. Taxa (superfamily, family, genus and species) have been updated according to current interpretations, based on reviews in the literature, the expertise of authors and contributors, and recently conducted fieldwork. Data to compile this catalogue were primarily gathered from the literature. In total, 1225 references were consulted and some information was also obtained from online databases. Each family was reviewed and the c…

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Revision of New World Species of Shore Fly Genus <I>Discomyza</I> Meigen (Diptera: Ephydridae)

Two species of Discomyza Meigen occur in the New World, one, D. u-signata Cresson, naturally and the other, D. maculipennis (Wiedemann), apparently as an introduction. These species are revised and their malacophagous breeding habits are documented. The species treatments include the first detailed descriptions and illustrations of structures from the male terminalia, lectotype designations for Notiphila maculipennis Wiedemann and its New World synonym (D. balioptera Loew), as well as phylogenetic and biogeographic information, including maps. Although congeners, these two species are apparently not closely related, and their occurrence in the New World is independent of each other. D. macu…

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