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Funding Information: We acknowledge the computational resources provided by the Aalto Science-IT project. We would like to thank Marita Kattelus, Tuomas Tolvanen and Toni Auranen from Advanced Magnetic Imaging (AMI) Centre for assistance in the neuroimaging measurements. We would like to thank Helge Kainulainen for providing us with engineering help. We would also like to thank Anni Nora, Nina Ritari, Laura Schildt and Ida ?rma performing the cognitive testing. Motion Analysis Laboratory team in Helsinki University Hospital measuring and analysing the sensorimotor testing, including Piia Haakana, Kati Tammivuori, Sari Korhonen, Elina Ylitalo, Tuula Niemel?, Essi Marttinen-Rossi, Juha-Pekka Kulmala and Jussi Nurminen. Funding Information: This work was supported by the Academy of Finland (grants #296240, #304294, #307250 and #3327288 to HP) and ”Brain changes across the life-span” profiling funding to University of Jyvaskyla (grant #311877).; Jane and Aatos Erkko Foundation and Emil Aaltonen Foundation (grant #602.274) to HP. Publisher Copyright: © 2021 The Author(s)